SPECIES OF EUCNEMIDAE
by Grabant, Aranka and Merkl, Ottó
contact: merkl_at_nhmus.hu
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This is a simple list of
Eucnemidae species housed in the Hungarian Natural History Museum,
without literature references. The real status of the types was not
studied, but it is almost certain that most of the "holotypes" and
"paratypes" ascribed to Reitter and other autors before 1950 are
actually syntypes.
HT = holotype, PT = paratype, LT = lectotype, PLT = paralectotype,
MT = "monotype" AT = allotype, CT = cotype
Year of description is unknown for us in a few cases; any help would be greatly acknowledged.
Species name Number of specimens Type included
Hungarian (more exactly: Carpathian Basin) material
Eucnemis capucina Ahrens, 1812: 34
Dromaeolus barnabita (Villa, 1838): 29
Thambus frivaldszkyi Bonvouloir, 1871: 13 HT
Microrhagus lepidus (Rosenhauer, 1847): 20
Microrhagus pygmaeus (Fabricius, 1792): 35
Microrhagus palmi Olexa, 1963: 3
Microrhagus emyi (Rouget, 1855): 32
Clypeorhagus clypeatus (Hampe, 1850): 8
Dirrhagofarsus attenuatus (Mäklin, 1848): 5
Rhacopus sahlbergi (Mannerheim, 1823): 10
Farsus dubius (Piller et Mitterpacher, 1783): 101
Hylis olexai (Palm, 1955): 24
Hylis cariniceps (Reitter, 1902): 4
Hylis simonae (Olexa, 1970): 1
Hylis foveicollis (Thomson, 1874): 100
Hylis procerulus (Mannerheim, 1823): 1
Xylophilus corticalis (Paykull, 1800): 22
Xylophilus testaceus (Herbst, 1806): 54 HT of Xylophilus flavipennis
Otho sphondyloides (Germar, 1818): 8
Melasis buprestoides (Linnaeus, 1761): 338
Isorhipis nigriceps (Mannerheim, 1823): 27
Isorhipis melasoides (Laporte, 1835): 143
Isorhipis marmottani (Bonvouloir, 1871): 29
Nematodes filum (Fabricius, 1801): 73
Palearctic material (excluding Carpathian Basin)
Eucnemis capucina Ahrens, 1812: 15
Dromaeolus barnabita Villa, 1838: 5
Dromaeolus maronita Bonvouloir, 1871: 1
Dromaeolus rufulus Fleutiaux, 1922: 1
Farsus dubius Piller et Mitterpacher,1783: 17
Dirrhagofarsus lewisi (Fleutiaux, 1901): 3
Dirhagus emyi Rouget, 1855: 4
Dirhagus ferrugineus Reitter, 1889: 2 HT, PT
Dirhagus hummleri Reitter, 1911: 1 HT
Dirhagus lepidus Rosenhauer, 1847: 4
Dirhagus pygmaeus Fabricius, 1792: 14
Rhacopus attenuatus Mäklin, 1845: 4 HT of Microrhagus longicornis
Rhacopus pyrenaeus Bonvouloir, 1872: 3
Rhacopus sahlbergi Mennerheim, 1823: 5
Isorhipis banghaasi Reitter, 1899 4 HT
Isorhipis melasoides Laporte, 1835: 27 HT, 2 PT of Isorhipis melasoides var. bicolor
Isorhipis nigriceps Mannerheim, 1823: 8
Isorhipis foveata Hisamatsu, 1955: 2
Melasis buprestoides Linnaeus, 1761: 20 HT, PT of Melasis buprestoides ab. semicastaneus
Melasis japonica Hisamatsu, 1963: 1
Anelastes druryi Kirby, 1818: 2
Hypocoelus cariniceps Reitter, 1902: 1
Hypocoelus foveicollis Thomson, 1874: 6
Hypocoelus olexai Palm, 1955: 17
Hypocoelus procerulus Mannerheim, 1823: 2 HT of Hypocoelus simplicifrons
Hypohilis matthiesseni Reitter, 1906: 3 HT, PT
Xylophilus corticalis Paykull, 1800: 8
Xylophilus humeralis Dufour, 1851: 1
Xylophilus seidlitzi Csiki, 1906: 1
Fornax nipponicus Fleutiaux, 1922: 1
Fornax lewisi Fleutiaux, 1922: 3
Xylophilus testaceus Herbst, 1806: 9
Hylochares cruentatus Gyllenhal, 1808: 3
Hylochares harmandi Fleutiaux, 1900: 1
Otho sphondyloides Germar, 1818: 6
Anelastidius feisthameli Graëlls, 1847: 3
Nematodes filum Fabricius, 1801: 5
Phyllocerus flavipennis Le Peletier de Saint-Fargeau et Audinet-Serville, 1825: 4
Non-Palearctic material
Cussolenis mutabilis Bonvouloir, 1875: 7 austr., orient.
Cussolenis robustus Fleutiaux, 1925: 1 afr.
Thylacosternus laetus Bonvouloir, 1871: 1 neotrop.
Lyssothyreus bimaculatus Laporte, 1835: 1 neotrop.
Lyssothyreus histrio Guérin-Méneville, 1844: 1 neotrop.
Gastraulacus bisulcatus Latreille, 1834: 2 : neotrop.
Eucalosoma versicolor Laporte, 1840 1 neotrop.
Pterotarsus albiventris Chevrolat, 1856: 2 orient
Pterotarsus auricolor Bonvouloir, 1875: 4 orient.
Pterotarsus bicolor Fleutiaux, 1856: 1 austr.
Pterotarsus bonvouloiri Fleutiaux, 1923: 2 austr.
Pterotarsus chevrolati Fleutiaux, 1923: 10 austr., orient.
Galbites chrysocoma (Hope, 1845): 2 orient.
Pterotarsus funebris Chevrolat, 1856: 6 orient., austr.
Pterotarsus marmoratus Guérin-Méneville, 1832: 10 austr.
Pterotarsus mouhoti Fleutiaux, 1928: 1 orient.
Pterotarsus niger Fleutiaux, 1897: 2 austr.
Pterotarsus ?tomentosus (Montrouzier, 1855): 1 orient.
Pterotarsus wallacei Perroud et Montrouzier, 1864: 12 orient., austr.
Galbites sedlaceki Muona, xxxx: 1 austr.
Galbites tigrina Muona, xxxx: 1 austr.
Galbites menieri Muona, xxxx: 1 austr.
Agastocerus confusus Fleutiaux, 1899: 4 orient.
Agastocerus frontalis Fleutiaux, 1899: 1 orient.
Vitellius lafertei Bonvouloir, 1875: 2 neotrop.
Poecilochrus intermedius Fleutiaux, 1896: 1 austr.
Submesogenus africanus Fleutiaux, 1899: 1 afr.
Euryostus reichei Bonvouloir, 1871: 1 orient.
Stethon pectorosus Leconte, 1866: 6 nearkt.
Deltometopus amoenicornis Say, 1836: 2 nearkt.
Galbidema mannerheimi Laporte, 1835: 1 austr.
Macroscython minimus Cobos, 1959: 2 afr.
Arisus Laportei Bonvouloir, 1871: 1 orient.
Arisus orientalis Laporte, 1835,: 1 orient.
Arisus adjunctus Bonvouloir, 1871: 1 austr.
Dendrocharis bicolor Redtenbacher, 1867: 1 orient.
Scopulifer atkinsoni Fleutiaux, 1912: 1 austr.
Scopulifer alternans (Bonvouloir, 1875): 1 orient.
Idiotarsus vestitus Bonvouloir, 1871: 1 neotrop.
Myall clavicornis (Lea, 1918): 1 austr.
Heterotaxis dubius Fleutiaux, 1896: 1 austr
Arganus distinctus Bonvouloir, 1871: 2 austr
Dromaeolus brevipes Fleutiaux, 1922: 6 orient., austr
Dromaeolus brevicornis Bonvouloir, 1871: 1 austr.
Dromaeolus javeti Bonvouloir, 1871: 4 neotrop.
Dromaeolus angustus Bonvouloir, 1871: 1 austr.
Dromaeolus lugubris Bonvouloir, 1871: 1 austr.
Dromaeolus opacus Bonvouloir, 1871: 1 orient.
Dromaeolus subcarinifrons Cobos, 1965: 1 afr.
Dromaeolus striatus (LeConte, 1852): 1 nearkt.
Dromaeolus cylindricollis (Say, 1836): 1 nearkt.
Dromaeolus semirugosus Fleutiaux, 1896: 2 afr.
Fornax adjectus Horn, 1890: 4 neotrop., orient.
Fornax cephalotes Fleutiaux, 1924: 1 orient.
Fornax concolor Blanchard, 1853: 308 orient., austr
Fornax incisus Bonvouloir, 1872: 2 orient.
Fornax infrequens Bonvouloir, 1872: 1 neotrop.
Fornax lineatus Bonvouloir, 1872: 1 austr
Fornax nicotianae Fleutiaux, 1895: 4 austr., orient.
Fornax orchesides Newman, 1838: 1 nearkt.
Fornax seponendus Bonvouloir, 1872: 3 orient.
Fornax sicardi Fleutiaux, xxxx: 1 afr.
Fornax variipennis Bonvouloir, 1872: 1 neotrop.
Fornax collega Bonvouloir, 1872: 2 orient.
Fornax substriatus Bonvouloir, 1872: 1 austr.
Fornax planus Bonvouloir, 1872: 8 austr., orient.
Fornax brunneus (Fleutiaux, 1896): 2 austr.
Fornax scythonoides Fleutiaux, xxxx: 1 afr.
Fornax brevis Fleutiaux, xxxx: 1 afr.
Fornax salax Bonvouloir, 1872: 1 neotrop.
Fornax pradieri (Bonvouloir, 1872): 1 afr.
Fornax fulvus Motschulsky, 1861: 1 orient.
Fornax bicolor (Melsheimer, 1846): 4 nearkt.
Fornax gabonensis Fleutiaux, 1896: 3 afr.
Onychodon canadensis (Brown, 1940): 2 nearkt.
Onychodon rugicollis (Fall, 1925): 2 nearkt.
Onychodon orchesides (Newman, 1838): 3 nearkt.
Isarthrus rufipes (Melsheimer, 1846): 1 nearkt.
Isarthrus calceatus (Say, 1836): 1 nearkt.
Asiocnemis hospitalis (Blanchard, 1904): 1 nearkt.
Thambus horni Muona, 2000: 1 nearkt.
Serrifornax tumidicollis (Redtenbacher, 1867): 1 orient.
Cyrtosthetus throscoides (Bonvouloir, 1872): 1 orient.
Sassanus colmanti Fleutiaux, 1925: 1 afr.
Chapianus monilicornis Fleutiaux, 1921: 1 orient.
Profornax languei Fleutiaux, 1896: 1 orient.
Fornax lacerdai Bonvouloir, 1872: 1 neotrop.
Fornax variepunctatus Bonvouloir, 1872: 1 neotrop.
Plesiofornax ligniperda Bonvouloir, 1872 1 neotrop.
Scython apicalis Bonvouloir, 1872: 2 orient.
Scython coloratum Bonvouloir, 1872: 1 orient.
Scython nigridorsus Cobos, xxxx rugidorsis 1965????: 2 afr.
Pseudoscython parvulus (Fleutiaux, 1899): 2 orient.
Farsus exoticus Bonvouloir, 1875: 1 austr.
Arrhipis albertisi Fleutiaux, 1896: 2 austr.
Arrhipis congoensis Cobos, 1969: 2 afr.
Arrhipis rugosa Fleutiaux, 1925: 26 afr.
Arrhipis senegalensis Laporte, 1835: 2 afr.
Arrhipis gaillardi Fleutiaux, xxxx: 5 afr.
Arrhipis sumatrensis Fleutiaux, 1897: 1 orient.
Arrhipis madagascariensis Fleutiaux, 1899: 2 afr.
Arrhipis orientalis Fleutiaux, 1896: 1 orient.
Arrhipis inimica Bonvouloir, 1875: 1 austr.
Arrhipis vassei Fleutiaux, xxxx: 3 afr.
Arrhipis horni Muona, xxxx: 1 neotr.
Arrhipis lanieri Guérin-Méneville, 1838: 2 nearkt., neortop.
Arrhipis subacuta Guérin-Méneville, 1843: 1 neotrop.
Sarfus spinosus Fleutiaux, 1925: 1 afr.
Entomophthalmus interruptus Bonvouloir, 1872: 1 neotrop.
Entomophthalmus rufiolus (LeConte, 1866): 1 nearkt.
Entomophthalmus africanus Fleutiaux, 1925 1 afr.
Dirhagus biroi Fleutiaux, 1898: 1 HT austr
Dirhagus triangularis Say, 1823: 2 nearkt.
Microrhagus crassus Bonvouloir, 1872: 1 orient.
Microrhagus derelictus Bonvouloir, 1872: 1 austr.
Microrhagus flabellatus Bonvouloir, 1872: 1 austr.
Microrhagus plaumanni (Cobos, 1965): 1 neotrop.
Microrhagus pectinatus LeConte, 1866: 1 nearkt
Prodirhagus dubius Fleutiaux, 1925: 1 afr.
Prodirhagus subparalleus (Bonvouloir, 1872): 1 orient.
Prodirhagus africanus (Schwarz, 1902): 1 afr.
Balistica vicina Fleutiaux, 1899: 1 orient.
Rhagomicrus humeralis Say,1836: 3 nearkt.
Rhagomicrus bonvouloiri (Horn, 1886): 1 nearkt.
Adelothyreus dufai Fleutiaux, 1907: 1 neotrop.
Arhagus bakeri Fleutiaux, 1921: 1 orient
Arhagus crassus Fleutiaux, 1935: 1 MT orient.
Ceratus albopilosus Fleutiaux, 1926: 1 orient.
Hylis terminalis (LeConte, 1866): 1 nearkt.
Golbachia impressicollis (Bonvouloir, 1872): 2 nearkt.
Isorhipis maculata Bonvouloir, 1875: 1 austr.
Isorhipis recticollis Fleutiaux, 1935: 1 MT austr.
Melasis rufipennis Horn, 1886: 17 nearkt.
Neomathion leprieuri Laporte, 1835: 1 neotrop.
Emathion foveicollis Bonvouloir, 1875: 1 neotrop.
Oisocerus murrayi Murray, 1863: 1 afr.
Euryptychus pasteuri (Fleutiaux, 1896): 1 orient.
Anelastes druryi Kirby, 1818: 9 nearkt.
Hylotastes ruficollis Bonvouloir, 1875: 2 orient.
Hemiopsida flavescens Bonvouloir, 1875: 1 austr
Hemiopsida nigricans Bonvouloir, 1875: 1 austr.
Hemiopsida nova Bonvouloir,1875: 1 austr.
Aubailius dercorsei Fleutiaux, 1921: 1 afr.
Curticephalus angustus Fleutiaux, 1912: 1 austr.
Nematodes guadeloupensis Fleutiaux,1911: 1 neotrop.
Nematodes suturalis Fleutiaux, 1896: 2 orient.
Nematodes infuscatus Bonvouloir, 1875: 1 neotrop.
Nematodes mexicanus Bonvouloir, 1835 1 neotrop.
Nematodes penetrans (LeConte, 1852): 1 nearkt.
Nematodes atropos (Say, 1836): 3 nearkt.
Hypocoelus nigricornis Say, 1823: 3 nearkt.
Fryanus gracilicornis Fleutiaux, 1935: 1 MT austr.
Mesogenus siamensis Fleutiaux, 1898: 1 orient.
Mesogenus harmondi Fleutiaux, 1898: 1 orient.
Mesogenus sumatrenis Fleutiaux, 1898: 1 orient.
Cryptosoma spinicorne Fabricius, 1801: 1 neotrop.
Potergus filiformis Bonvouloir, 1871: 3 orient.
Epiphanis cornutus Eschscholtz, 1829: 1 nearkt.
Perothops witticki LeConte, 1857: 1 nearkt.
Isorhipis obliqua (Say, 1836): 10 nearkt.
Isorhipis ruficornis (Say, 1823): 10 nearkt.
Xylophilus orientalis (Fleutiaux, 1896): 1 orient.
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2009. Hungarian Natural History Museum,
Department of Zoology, Coleoptera Collection